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  • BSc (Hons) Biological Sciences, Imperial College, London
  • MSc Palaeobiology, University of Bristol
  • PhD The anatomy and biomechanics of elephant limbs and feet, supervised by Prof. John Hutchinson, The Royal Veterinary College, University of London
  • PGDPPHE, University of South Wales
  • Senior Fellow of The Higher Education Academy


Dr Charlotte Miller is a comparative anatomist. At the University of South Wales she teaches vertebrate structure and function, including humans, with particular specialisms in limbs and locomotion.

Charlotte’s experience includes work with medical (Duke University, USA) and veterinary (The Royal Veterinary College) anatomy. For her PhD she studied elephant limb anatomy and function, and her masters research looked at bird claw shape as a potential method for reconstructing dinosaur behaviour.


Modules lead:

  • Human Anatomy and Physiology (Year 1 Human Biology and Medical Science)
  • Human Form and Function (Year 2 Human Biology and Medical Science)
  • Vertebrate Zoology (Year 2 Biology, International Wildlife Biology, Natural History)
  • Human Structure (Year 3 Medical Science)

Module contributions:

  • Clinical/Applied Physiology (Year 2 Human Biology and Medical Science)
  • Frontiers in Biology (Year 2 Biology)


  • Griffin, N.L., Miller, C.E., Schmitt, D., D’Aout, K (2015) Understanding the evolution of the windlass mechanism of the human foot from comparative anatomy: insights, obstacles, and future directions. American Journal of Physical Anthropolgy 156(1):1-10
  • Wunderlich, R.E., Tongen, A., Gardiner, J., Miller, C.E., Schmitt, D. (2014) Dynamics of Locomotor Transitions from Arboreal to Terresttrial Substrates in Verraux’s Sifaka (Propithecus verreauxi). Integrative and Comparative Biology. Sep 17. pii: icu110. [Epub ahead of print]
  • Granatosky, M.C., Miller, C.E., Boyer, D.M., Schmitt, D. (2014) Lumbar vertebral morphology of flying, gliding and suspensory mammals: Implications for the locomotor behavior of the subfossil lemurs Palaeopropithecus and Babakotia. Journal of Human Evolution, 75:40-52
  • Sparling, T.L., Schmitt, D., Miller, C.E., Guilak, F., Somers, T.J., Keefe, F.J., Queen, R.M. (2014) Energy Recovery in Individuals with Knee Osteoarthritis. Osteoarthritis and Cartilage. DOI: 22(6):747-55
  • Nicole L Griffin, Charlotte Miller, Daniel Schmitt, Kristiaan D’Août (2013) An investigation of the dynamic relationship between navicular drop and first metatarsophalangeal joint dorsal excursion. Journal of Anatomy, 222(6):571-644.
  • Aleksandra V Birn-Jeffery, Charlotte E Miller, Darren Naish, Emily J Rayfield, David W E Hone (2012) Pedal claw curvature in birds, lizards and mesozoic dinosaurs – complicated categories and compensating for mass-specific and phylogenetic control. PloS ONE, 7(12):e50555.
  • John R Hutchinson, Cyrille Delmer, Charlotte E Miller, Thomas Hildebrandt, Andrew A Pitsillides, Alan Boyde (2011) From flat foot to fat foot: structure, ontogeny, function, and evolution of elephant “sixth toes”. Science, 334(6063):1699-703.
  • Lei Ren, Charlotte E Miller, Richard Lair, John R Hutchinson (2010) Integration of biomechanical compliance, leverage, and power in elephant limbs. PNAS, 107(15):7078-82.
  • Lei Ren, Melanie Butler, Charlotte Miller, Heather Paxton, Delf Schwerda, Martin S Fischer, John R Hutchinson (2008) The movements of limb segments and joints during locomotion in African and Asian elephants. Journal of Experimental Biology, 211(17):2735-51.
  • Charlotte E Miller, Christopher Basu, Guido Fritsch, Thomas Hildebrandt, John R Hutchinson (2008) Ontogenetic scaling of foot musculoskeletal anatomy in elephants. Journal of the Royal Society, Interface. 5(21):465-75.
  • John R Hutchinson, Charlotte E Miller, Guido Fritsch, Thomas Hildebrandt (2008) The anatomical foundation for multidisciplinary studies of animal limb function: examples from dinosaur and elephant limb imaging studies. pp. 23-38 in H. Endo and R. Frey (eds.), Anatomical Imaging Techniques: Towards a New Morphology. Berlin: Springer-Verlag.

Research Gate profile


International society for vertebrate morphology


Charlotte’s research concentrates on animal:substrate interactions, particularly moving safely across a surface without falling or getting injured. Most recently she has been working on primate movement with collaborators in the Animal Locomotion Lab at Duke University, James Madison University, and the University of Alberta.

Research techniques:

  • Force plate, video, image processing, motion analysis and accelerometry techniques for studying locomotion, including slow and fast gaits, leaping and landing.
  • MATLAB programming.
  • Reconstruction of internal anatomy from Medical imaging data, and anatomical dissection, including muscle architecture and moment arm analysis.


@Dr_Charlotte_M @USW_Biology

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