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B.Sc. (Hons), Ph.D.


2011 – present. Reader of Mathematics, University of South Wales

2003-2011. Lecturer/Senior Lecturer in Mathematical Sciences, University of Glamorgan.

2000-2003. Postdoctoral Research Assistant, Division of Mathematics, University of Dundee (employed on BBSRC 94/MAF12243).

1996-1999. PhD Student, Department of Mathematics, University of Bath. Employed part-time as Graduate Teaching Assistant.

In 2004 I was awarded a Nuffield Foundation grant for Newly Appointed Lecturers in Science, Engineering and Mathematics (NUF-NAL 04) to investigate biobarriers as tools for bioremediation.


Main Teaching subjects and course leadership:
Nonlinear Systems and Mathematical Biology
Linear Algebra

Main Administrative Activities
Unit of Assessment leader for Mathematical Sciences in REF2014
Undergraduate admissions tutor for Mathematics


Selected publications include:

Hayward J. & Boswell G.P. (2014)
Model Behaviour and the Concept of Loop Impact: A Practical Method. System Dynamics Review, 30(1), 29-57

Boswell G.P. (2012)
Modelling combat strategies in fungal mycelia. Journal of Theoretical Biology. 304: 226-234.

Boswell G.P. & Davidson F.A. (2012)
Modelling hyphal networks. Fungal Biology Reviews, 26(1):30-38.

Boswell G.P. (2012)
Power variation strategies for cycling time trials: a differential equation model. Journal of Sports Sciences, 30(7): 651-659.

Hopkins S. & Boswell G.P. (2012)
Mycelial response to spatiotemporal nutrient heterogeneity: a velocity-jump mathematical model. Fungal Ecology 5(2): 124-136.

Davidson F.A., Boswell G.P., Fischer M.W.F., Heaton L., Hofstadler D. and Roper M. (2011)
Mathematical modelling of fungal growth and function IMA Fungus 2(1): 33-37.

Boswell G.P. (2008)
Modelling mycelial networks in structured environments. Mycological Research, 112: 1015-1025.

Boswell G.P. and Hopkins S. (2008)
Linking hyphal growth to colony dynamics: spatially explicit models of mycelia. Fungal Ecology, 1(4): 143-154.

Carver I. and Boswell G.P. (2008)
A lattice-free model of translocation induced outgrowth in mycelial fungi. IAENG International Journal of Applied Mathematics 38(4): 173-179.

Boswell G.P. & Davidson F.A. (2007).
Diffusion fronts in enzyme-catalysed reactions. Journal of Engineering Mathematics, 59: 157-169.

Boswell G.P., Jacobs H., Ritz K., Gadd G.M. & Davidson F.A. (2007).
The development of fungal networks in complex environments. Bulletin of Mathematical Biology, 69: 605-634.

Jacobs H., Boswell G. P., Scrimgeour C. M., Davidson F. A., Gadd G. M. & Ritz K. (2004)
Translocation of carbon by Rhizoctonia solani in nutritionally-heterogeneous environments. Mycological Research, 108: 453-462.

Boswell G.P., Jacobs H., Gadd G.M., Ritz K. & Davidson F.A. (2003).
A mathematical approach to studying fungal mycelia. Mycologist, 17, 165-175.

Boswell G.P., Jacobs H., Davidson F.A., Gadd G.M. & Ritz K. (2003).
Growth and function of fungal mycelia in heterogeneous environments. Bulletin of Mathematical Biology, 65, 447-477.

Boswell, G.P., Jacobs H., Davidson F.A., Gadd G.M. & Ritz K. (2003).
A positive numerical scheme applied to a mixed-type 1D partial differential equation model for fungal growth. Applied Mathematics and Computation, 138, 321-340.

Boswell, G.P., Jacobs H., Davidson F.A., Gadd G.M. & Ritz K. (2002).
Functional consequences of nutrient translocation in mycelial fungi. Journal of Theoretical Biology, 217, 459-477.

Jacobs, H., Boswell G. P., Ritz K., Davidson F. A. & Gadd G. M. (2002)
Solubilization of calcium phosphate as a functional consequence of glucose translocation by Rhizoctonia solani.
FEMS Microbiology Ecology, 40: 65-71.

Jacobs, H., Boswell G. P., Harper F. A., Ritz K., Davidson F. A. & Gadd G. M. (2002)
Solubilization of metal phosphates by Rhizoctonia solani.
Mycological Research, 106: 1468-1479.

Boswell G.P., Franks N.R. & Britton N.F. (2001).
Arms Races and the Evolution of Big Fierce Societies. Proceedings of the Royal Society of London, B, Biological Sciences, 268, 1723-1730.

Britton, NF, Boswell, GP & Franks, NR. (2001) Dispersal and conservation in fragmented landscapes Insect Movement: Mechanisms and Consequences , Proceedings of the 20th Symposium of the Royal Entomological Society. CAB International , 299 – 320.

Boswell G.P., Franks N.R. & Britton N.F. (2000).
Habitat fragmentation and swarm raiding army ants. In Behavior and Conservation (Eds. Gosling L.M. & Sutherland W.J.). Cambridge University Press, Cambridge. pp. 141-158.

Boswell G.P., Britton N.F. & Franks N.R. (1998).
Habitat fragmentation, percolation theory and the conservation of a key-stone species. Proceedings of the Royal Society of London, B, Biological Sciences, 265, 1921-1925.


Member of the Society of Mathematical Biology (SMB).


Member of Research Committee of the Welsh Institute of Mathematical and Computational Sciences.

Member of Scientific Committee for BAMC 2014

Member of the Society of Mathematical Biology.

Member of Editorial Board for Network Biology


My research is primarily in mathematical biology, with particular emphasis on ecological applications. Past and current projects include modelling:

Habitat fragmentation for E.burchilli

The evolution of gigantism in societies

The impact of heterogeneous growth conditions for fungi using PDEs, lattice-based and lattice-free approaches

Enzyme-kinetics and Michaelis-Menton theory

Biobarriers as tools for bioremediation

Cycling performance

Mathematical modelling of whole-body bio-energtic processes with applications in sports

I have successfully co-supervised 2 PhD student (models of bilingualism and lattice-free models of mycelial networks).

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